Name: Capnocytophaga granulosa Yamamoto et al. 1994

Category: Species

Proposed as: sp. nov.

Etymology: gra.nu.lo’sa. L. neut. dim. n. granulum, a small grain; L. fem. adj. suff. -osa, suffix meaning full of; N.L. fem. adj. granulosa, granular

Pronunciation, gender: gra-nu-LO-sa, feminine

Type strain: ATCC 51502; B 611; CCUG 32991; CIP 104128; DSM 11449; JCM 8566; LMG 16022; NCTC 12948

See detailed strain information at

Conduct genome-based taxonomy at

16S rRNA gene: X97248 Analyse FASTA

Valid publication: Yamamoto T, Kajiura S, Hirai Y, Watanabe T. Capnocytophaga haemolytica sp. nov. and Capnocytophaga granulosa sp. nov., from human dental plaque. Int J Syst Bacteriol 1994; 44:324-329.

IJSEM list: Anonymous. Notification list. Notification that new names and new combinations have appeared in volume 44, part 2 of the IJSB. Int J Syst Bacteriol 1994; 44:596.

Nomenclatural status: validly published under the ICNP

Taxonomic status: correct name (and explicitly recommended for medical use)

Risk group: 3

• Notes:
  • 😷 The risk group for Belgium has been imported on 2024-02-05. The full classification is: risk group = 2, note = "human pathogen". — The risk group for Canada has been imported on 2024-02-27. The full classification is: risk group = 2, note = "Animal classification RG: 1 - Security sensitive biological agent: No - Terrestrial animal pathogen under Canadian Food Inspection Agency authority: No - Containment level: Containment Level 2". — The risk group for Swiss Confederation has been imported on 2024-02-01. The full classification is: risk group = 3, note = "TMP". — The risk group for Germany has been imported on 2023-10-29. The full classification is: risk group = 2. — If in doubt, use the risk group given in the regulations for your country and, if these are not available, use the risk group given in the catalogue of the culture collection from which you have obtained or intend to obtain the strain.
  • 🎓 Name mentioned 14 times in PubMed until 2025-06-04.
  • 🙆 This name is on the List of Recommended Names for bacteria of medical importance (LoRN) as the name to be applied to this species. The name is on LoRN because of the recorded risk group of the species itself.
  • 🧍 According to Bartlett et al. (2022), this species is an established human pathogen. Publication: Bartlett A, Padfield D, Lear L, Bendall R, Vos M. A comprehensive list of bacterial pathogens infecting humans. Microbiology (Reading) 2022; 168:0.
  • 🧍 The genome of the type strain was sequenced as part of the GEBA (Genomic Encyclopedia of Bacteria and Archaea) project. Publication: Garcia-Lopez M, Meier-Kolthoff JP, Tindall BJ, Gronow S, Woyke T, Kyrpides NC, Hahnke RL, Goker M. Analysis of 1,000 Type-Strain Genomes Improves Taxonomic Classification of Bacteroidetes. Front Microbiol 2019; 10:2083.
  • 🧍 The phylogenetic position of this species was determined in a genome-scale analysis by Hahnke et al. (2016). Publication: Hahnke RL, Meier-Kolthoff JP, Garcia-Lopez M, Mukherjee S, Huntemann M, Ivanova NN, Woyke T, Kyrpides NC, Klenk HP, Goker M. Genome-Based Taxonomic Classification of Bacteroidetes. Front Microbiol 2016; 7:2003.

Parent taxon: Capnocytophaga Leadbetter et al. 1982

Assigned by: Yamamoto T, Kajiura S, Hirai Y, Watanabe T. Capnocytophaga haemolytica sp. nov. and Capnocytophaga granulosa sp. nov., from human dental plaque. Int J Syst Bacteriol 1994; 44:324-329.

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Record number: 774467