Species names derived from present participles A-M

General remarks

Species epithets whose last component is a present participle (usually ending in -ens or -ans) can be in any gender. They are treated like Group B adjectives and belong to the third declension. The actual gender is determined by the gender of the genus name [Rule 12c (1)]. One part of the list of epithets formed in that manner is provided below. Genus names and subspecies names derived from present participles are not listed here. A special case is genes, which originally is an Ancient Greek suffix meaning "born from" but is nowadays mainly used as a Neolatin suffix meaning "producing". See also the LPSN introductory page on forming names.

New combinations

In the case of a new combination these epithets must not be adapted to the gender of the genus name. This is a special case of Group B adjectives.

brunescens

Lysobacter brunescens

canescens

Streptomyces canescens

captans

Gluconacetobacter azotocaptans

cellulans

Cellulomonas cellulansCellulosimicrobium cellulansNocardia cellulans

circulans

Bacillus circulansNiallia circulans

coagulans

Bacillus coagulansBacteroides coagulansBombiscardovia coagulansEzakiella coagulansHyphomicrobium coagulansStaphylococcus coagulansWeizmannia coagulans

coalescens

Hymenobacter coalescensThermococcus coalescens

coelescens

"Actinomyces coelescens"Streptomyces coelescens

coerulescens

"Actinomyces coerulescens"Bacularia coerulescensStreptomyces coerulescens

cohabitans

Rectinema cohabitans

cohaerens

Herpetosiphon cohaerensLampropedia cohaerensLewinella cohaerens

colans

Thermococcus fumicolans

colens

Achromobacter mucicolensActinomyces denticolensAlloscardovia omnicolensBifidobacterium denticolensCorynebacterium accolensPaenibacillus planticolensPandoraea horticolensParascardovia denticolensPedobacter planticolensPyramidobacter piscolensTreponema isoptericolens

comitans

Actinobacillus actinomycetemcomitansAggregatibacter actinomycetemcomitans"Bacterium actinomycetemcomitans"Haemophilus actinomycetemcomitansRoseivirga echinicomitansVibrio comitans

commovens

Pandoraea commovens

congelans

Pseudomonas congelans

contaminans

Alicyclobacillus contaminansAnoxybacillus contaminansBurkholderia contaminansChryseobacterium contaminansHerminiimonas contaminansLysinibacillus contaminansOrnithinibacillus contaminansPaenibacillus contaminansParacoccus contaminansPseudochelatococcus contaminansRahnella contaminansSphingopyxis contaminans"Sphingorhabdus contaminans"Sphingorhabdus contaminansSporosarcina contaminans

convertens

Slackia isoflavoniconvertensSphingobium aromaticiconvertens

corrodens

"Bacteroides corrodens"Eikenella corrodens

crescens

Aquimarina celericrescensArthrobacter celericrescensBacillus crescensClostridium celerecrescensCohaesibacter celericrescensFlavobacterium celericrescensKluyvera cryocrescensLacrimispora celerecrescensLiberibacter crescensMarinobacter segnicrescensMethylophaga nitratireducenticrescensOryzihumus leptocrescensRobertmurraya crescensSpiroplasma velocicrescensThermococcus celericrescensThiobacillus rapidicrescens

cumulans

Acidiplasma cupricumulansAcinetobacter cumulansCatenisphaera adipataccumulansFerroplasma cupricumulansVariovorax boronicumulans

defendens

Frateuria defendens

defragrans

Alcaligenes defragransCastellaniella defragrans

degradans

"Microbulbifer degradans"Olivibacter oleidegradansRhodococcus degradansSaccharophagus degradans

delens

Nocardia cummidelensNonlabens xylanidelensPersicivirga xylanidelens

deligens

Williamsia deligens

desmolans

Agathobaculum desmolansButyricicoccus desmolansEubacterium desmolans

detergens

Sphingobacterium detergens

devorans

"Bacillus devorans"Flavobacterium devoransPaenibacillus agaridevorans"Steroidobacter agaridevorans"Thalassobacillus devorans

diffluens

Cytophaga diffluensPersicobacter diffluens

disiens

Bacteroides disiensPrevotella disiens

divergens

Carnobacterium divergensLactobacillus divergens

dolens

Achromobacter dolensPseudomonas mucidolens

durans

Acidithiobacillus ferriduransAcidocella aluminiiduransAdhaeribacter radioduransAlcalilimnicola haloduransAlkalihalobacillus haloduransAlkalilimnicola halodurans corrig.Alsobacter metallidurans"Amycolatopsis acididurans"Aridibacter famiduransArthrobacter haloduransBacillus haloduransBacillus psychroduransBacillus saliduransBacillus sporothermoduransCompanilactobacillus haloduransCupriavidus metalliduransDeinococcus radioduransDesulfosporosinus acididuransDyella haloduransDyella psychroduransEnterococcus duransFerdinandcohnia saliduransFlavobacterium aciduransFredinandcohnia salidurans corrig."Candidatus Gallionella acididurans"Halomonas haloduransHalomonas zinciduransHalorubrum haloduransHeyndrickxia sporothermodurans"Hymenobacter radiodurans"Lactobacillus haloduransLentibacillus haloduransMaribacter cobaltiduransMarinobacter haloduransMarivibrio haloduransMesorhizobium metalliduransMethanobrevibacter acididuransMethylobacterium durans"Methylobacterium radiodurans"Microbacterium radioduransMicromonospora nickeliduransNocardioides euryhaloduransPaenibacillus xerothermoduransPediococcus ethanoliduransPsychrobacillus psychroduransRalstonia metalliduransRhizobium metalliduransRoseivivax haloduransRoseomonas radioduransRoseomonas selenitiduransSalinicoccus haloduransStreptococcus duransStreptomyces acididuransThalassococcus haloduransWautersia metallidurans

edens

Burkholderia peredensCaballeronia peredensDefluviitalea raffinosedensHalorubrum glutamatedensPseudomonas nicosulfuronedensWilliamsia serinedens

efficiens

Bradyrhizobium diazoefficiensCorynebacterium efficiens

egens

Altererythrobacter salegensArenibacter catalasegensArthrobacter terregens"Ascidiaceibacter salegens"Aureobacterium terregensBeneckea natriegensCroceibacterium salegensFlavobacterium salegensHallella seregens"Leptotrichia sanguinegens"Microbacterium terregensPseudidiomarina gelatinasegens"Pseudomonas natriegens"Pseudomonas protegensPseudomonas salegensSalegentibacter salegensSneathia sanguinegensVibrio natriegens

elabens

Coccochloris elabens"Micraloa elabens"Microcystis elabens

eligens

Eubacterium eligensLachnospira eligens

elongans

Haloferax elongans

eludens

"Candidatus Ruthmannia eludens"

erumpens

Streptomyces erumpens

exalbescens

Nesiotobacter exalbescensPseudovibrio exalbescens

excellens

Marinobacter excellens

exedens

Advenella alkanexedens"Arenimonas alkanexedens"Dehalogenimonas formicexedensDesulfoglaeba alkanexedensNannocystis exedensPaenibacillus agarexedensPaenibacillus xylanexedens

exhalans

Nocardiopsis exhalans

exigens

Aestuariibacter salexigensBacillus salexigensBryantella formatexigensChromatium salexigensChromohalobacter salexigensDesulfocapsa sulfexigens corrig.Desulfovibrio salexigensDesulfuromonas acetexigensGramella salexigensHaliea salexigensHalochromatium salexigensMaridesulfovibrio salexigensMarinobacter salexigensMarinobacter salinexigensMarvinbryantia formatexigensMelitea salexigensMycoavidus cysteinexigensPararhodospirillum sulfurexigensRheinheimera salexigens"Rheinheimera salexigens"Rhodopseudomonas pentothenatexigensRhodospirillum salexigensRhodospirillum sulfurexigensRhodothalassium salexigensSalibacillus salexigensSalimicrobium salexigensThalassobaculum salexigens"Trichloromonas acetexigens"Virgibacillus salexigens

extorquens

Methylobacterium extorquensMethylorubrum extorquensProtomonas extorquens

faciens

Acetobacter liquefaciensAcetobacter liquefaciensAchromobacter aegrifaciensAcidisphaera rubrifaciensAdlercreutzia equolifaciens"Aerobacter liquefaciens"Agrobacterium tumefaciensAlteromonas nigrifaciensAlteromonas putrefaciensAquibacter zeaxanthinifaciensArthrobacter ureafaciensAureobacterium liquefaciensBacillus acidinfaciensBacillus amyloliquefaciensBacillus pulvifaciens"Bacillus putrefaciens""Bacterium flaccumfaciens""Bacterium tumefaciens"Bacteroides acidifaciens"Bacteroides aerofaciens"Bacteroides propionicifaciensBrenneria rubrifaciensBrevibacterium liquefaciensClostridium putrefaciensCollinsella aerofaciensCorynebacterium flaccumfaciensCorynebacterium liquefaciensCorynebacterium mucifaciens"Corynebacterium ureafaciens"Cosenzaea myxofaciensCurtobacterium flaccumfaciensDialister propionicifaciensDyella amyloliquefaciens"Eberthella alcalifaciens"Ellagibacter isourolithinifaciens"Candidatus Endobryopsis kahalalidifaciens" corrig.Erwinia rubrifaciensEubacterium aerofaciens"Ferrovum myxofaciens"Globicatella sulfidifaciensGluconacetobacter liquefaciens corrig.Gluconoacetobacter liquefaciens"Gluconobacter liquefaciens" corrig.Gordonibacter urolithinfaciensKitasatospora aureofaciensLactobacillus kefiranofaciensLentibacillus amyloliquefaciensMesoflavibacter zeaxanthinifaciensMicrobacterium liquefaciensMicromonospora viridifaciensMoraxella nonliquefaciensMycoplasma lipofaciensMycoplasma putrefaciensMycoplasmopsis lipofaciensNatronorubrum sulfidifaciensNubsella zeaxanthinifaciensPaenarthrobacter ureafaciensPaenibacillus pulvifaciensParabacteroides acidifaciensParacoccus carotinifaciensParacoccus zeaxanthinifaciens"Pedobacter zeaxanthinifaciens"Propionibacterium acidifaciensProteus myxofaciensProvidencia alcalifaciensPseudoalteromonas nigrifaciensPseudomonas aureofaciensRhizomonas suberifaciensRhizorhapis suberifaciensRobiginitalea myxolifaciensRubritalea squalenifaciensRuminococcus flavefaciensSalinicoccus halitifaciensSerratia liquefaciensShewanella putrefaciensSiansivirga zeaxanthinifaciensSiminovitchia acidinfaciensSlackia equolifaciensSphingobium suberifaciensSphingomicrobium astaxanthinifaciensSphingomonas astaxanthinifaciensSphingomonas carotinifaciensSphingomonas floccifaciensSphingomonas spermidinifaciensSphingomonas suberifaciensStreptomyces albofaciensStreptomyces ambofaciensStreptomyces aureofaciensStreptomyces graminofaciensStreptomyces melanosporofaciensStreptomyces nigrifaciensStreptomyces phaeofaciensStreptomyces roseifaciensStreptomyces sulfonofaciensSulfobacillus benefaciensThermus amyloliquefaciensTreponema succinifaciens

fascians

Corynebacterium fascians"Phytomonas fascians"Rhodococcus fascians

fermentans

"Abyssisolibacter fermentans"Acetobacter oryzifermentansAcidaminococcus fermentansActinotalea fermentansAlkalibacter saccharofermentansAlkaliphilus peptidifermentans corrig.Anaerotignum lactatifermentansAnoxybacter fermentans"Bacillus bifermentans"Bacillus glycinifermentansBrachybacterium tyrofermentans"Brevefilum fermentans"Brevibacterium fermentans"Candidatus Caldatribacterium saccharofermentans"Caproicibacter fermentansCellulomonas fermentansClostridium bifermentans"Clostridium cellulofermentans"Clostridium lactatifermentansClostridium phytofermentansCorynebacterium afermentansCytophaga fermentansDesulfurococcus fermentansDyadobacter fermentansGallibacterium trehalosifermentansGeothrix fermentansGluconacetobacter sucrofermentansHalanaerobium fermentans corrig.Haloanaerobium fermentans"Komagatabacter sucrofermentans"Komagataeibacter sucrofermentans"Lachnoclostridium phytofermentans"Lactiplantibacillus fabifermentansLactobacillus bifermentansLactobacillus fabifermentansLactobacillus malefermentansLactobacillus oligofermentansLoigolactobacillus bifermentansMacellibacteroides fermentansMycoplasma fermentansMycoplasmopsis fermentansPaludibaculum fermentansParaclostridium bifermentansPaucilactobacillus oligofermentans"Pelorhabdus rhamnosifermentans"Pelosinus fermentansProteiniphilum saccharofermentans"Candidatus Pseudoramibacter fermentans""Psychrosinus fermentans"Rhodoferax fermentansRodentibacter trehalosifermentansRoseiarcus fermentansRuminiclostridium herbifermentansSaccharicrinis fermentansSecundilactobacillus malefermentansStaphylococcus piscifermentansStreptococcus oligofermentansTepidibacillus fermentansVagococcus acidifermentans

fervens

Methanocaldococcus fervensMethanococcus fervensStreptomyces fervens"Streptomyces fervens"Streptoverticillium fervens

figens

Acetobacter nitrogenifigensAhniella affigensCiceribacter azotifigensFlavobacterium nitrogenifigensIdeonella azotifigensNatronobacillus azotifigensNovosphingobium nitrogenifigensOleiliquidispirillum nitrogeniifigensPaenibacillus azotifigensParaburkholderia azotifigensPseudomonas azotifigensSphingomonas azotifigens

flagrans

Arvibacter flaviflagransFlavobacterium flaviflagrans

flavescens

Arthrobacter flavescensAureobacterium flavescensConservatibacter flavescensCorynebacterium flavescensEnterococcus flavescens"Flavobacterium flavescens"Leptobacterium flavescensLuteolibacter flavescensMarinomonas flavescensMethanosarcina flavescensMicrobacterium flavescensMicrovirga flavescensMuricauda flavescensMycobacterium flavescensMycolicibacterium flavescensMyxococcus flavescensNeisseria flavescensNocardioides flavescensNocardiopsis flavescensPseudomonas flavescens"Rhizobium flavescens"Streptomyces albiflavescensTessaracoccus flavescensVirgibacillus flavescens

flectens

Phaseolibacter flectensPseudomonas flectens

flocculans

Balneimonas flocculans corrig.Balneomonas flocculans"Comamonas flocculans"Hymenobacter flocculansMicrovirga flocculansRubritepida flocculans"Siccationidurans flocculans"Sinimarinibacterium flocculans

fluorescens

Actinomadura luteofluorescens"Actinomyces citreofluorescens""Actinomyces fluorescens"Pseudomonas fluorescensStreptomyces citreofluorescensStreptomyces fluorescens"Streptomyces luteofluorescens"

formans

Acetomicrobium hydrogeniformansAcidaminobacter hydrogenoformansActinomadura sporangiiformansAllisonella histaminiformansAnaerobaculum hydrogeniformansBacillus azotoformansBacillus ectoiniformans"Bacillus gonidiaformans"Bacteroides zoogleoformans"Caecibacterium sporoformans"Caecibacterium sporoformansCalidifontibacillus azotoformansCapsularis zoogleoformans corrig.Carboxydothermus hydrogenoformansCereibacter azotoformansClostridium hydrogeniformans"Corynebacterium laevaniformans"Dulcicalothrix necridiiformansFaecalicoccus acidiformansFortiea necridiiformansFusobacterium gonidiaformans"Halanaerobium hydrogenoformans"Luteovulum azotoformans"Luteovulum azotoformans""Candidatus Manganitrophus noduliformans"Meiothermus cateniformansMicrobacterium laevaniformans"Microbispora cellulosiformans"Nocardiopsis synnemataformans"Peteryoungia rosettiformans"Prevotella zoogleoformansProteinivorax hydrogeniformansPseudomonas azotoformansRhizobium rosettiformansRhodobacter azotoformansRuthenibacterium lactatiformansSchinkia azotoformansStreptomyces sporangiiformansStreptomyces synnematoformansSuicoccus acidiformansThermoanaerobacterium butyriciformansThermosipho globiformans

frangens

Nocardia pigrifrangens

frigens

Deinococcus frigens

fugiens

"Candidatus Haliscomenobacter calcifugiens"

fulvescens

Actinomadura fulvescens

furvescens

Marinoscillum furvescens

fuscans

Streptacidiphilus fuscansXanthomonas fuscans

genens

Alkalispirochaeta sphaeroplastigenensMechercharimyces asporophorigenensSphingomonas sanxanigenensSpirochaeta sphaeroplastigenensVirgisporangium aliadipatigenens

generans

Dorea formicigeneransEubacterium formicigenerans

-genes (genes)

Acetobacteroides hydrogenigenesActinobacillus succinogenesActinomadura nitritigenes"Actinomyces diastatochromogenes"Actinomyces pyogenesActinomyces pyogenes"Actinomyces violaceochromogenes""Actinomyces viridochromogenes"Actinoplanes ianthinogenes"Acuformis alcaligenes"Aeromonas enteropelogenesAgrobacterium rhizogenesAlkalibaculum sporogenes"Anaerophilus nitritogenes"Aquitalea pelogenesArcanobacterium pyogenesArthrobacter polychromogenesAzomonas macrocytogenesAzomonotrichon macrocytogenesAzotobacter macrocytogenes"Bacillus necrogenes""Bacillus pyogenes""Bacillus scatologenes""Bacillus sporogenes""Bacterium ammoniagenes""Bacterium monocytogenes"Bacteroides helcogenesBacteroides pyogenesBacteroides succinogenesBeneckea gazogenesBiomaibacter acetigenesBrevibacterium ammoniagenesCaminicella sporogenesChryseobacterium indologenesClostridium gasigenesClostridium nitritogenesClostridium scatologenesClostridium sporogenesClostridium sulfidigenesClostridium thermosuccinogenesClostridium thermosulfurigenes corrig."Colobactrum aerogenes"Corynebacterium ammoniagenesCorynebacterium pyogenesCurtobacterium ammoniigenesDesulfocapsa thiozymogenesDesulfonispora thiosulfatigenesEnterobacter aerogenesErcella succinigenesEubacterium coprostanoligenesEubacterium nitritogenesFibrobacter succinogenesFlavobacterium indologenesFusobacterium necrogenesGeobacter thiogenes"Hallerella succinigenes"Halopelagius fulvigenesHaloplanus aerogenes"Helcococcus pyogenes"Kingella indologenesKlebsiella aerogenesLechevalieria aerocolonigenesLentzea aerocolonigenes"Leptothrix trichogenes"Listeria monocytogenesLysobacter enzymogenesMediterraneibacter butyricigenesMethylobacillus glycogenesMicrobispora amethystogenesMicrobispora chromogenesMicromonospora purpureochromogenesMoryella indoligenesMyceligenerans crystallogenesMycobacterium farcinogenesMycolicibacterium farcinogenes"Necromonas achromogenes""Nitrosopumilus cobalaminigenes""Nocardia aerocolonigenes"Nocardiopsis chromatogenesOlsenella scatoligenesOscillibacter valericigenesOxalobacter formigenesPaludibacter propionicigenesPasteurella aerogenesPelobacter seleniigenes"Phytomonas rhizogenes"Proteiniborus ethanoligenesProteiniphilum acetatigenesPseudarthrobacter polychromogenes"Pseudoclostridium thermosuccinogenes"Pseudomonas alcaligenesPseudomonas pseudoalcaligenesPsychrobacter fulvigenesPyrinomonas methylaliphatogenesRhizobium rhizogenesRhizobium tumorigenesSaccharofermentans acetigenesSaccharothrix aerocolonigenesSeleniibacterium seleniigenesSporanaerobacter acetigenesStaphylococcus chromogenesStreptacidiphilus melanogenesStreptococcus pyogenesStreptomyces achromogenesStreptomyces albidochromogenesStreptomyces cinerochromogenesStreptomyces cyaneochromogenesStreptomyces diastatochromogenesStreptomyces fuscichromogenesStreptomyces fuscigenesStreptomyces griseochromogenesStreptomyces indoligenesStreptomyces melanogenesStreptomyces olivochromogenesStreptomyces parvisporogenesStreptomyces phaeochromogenesStreptomyces phaeogriseichromatogenesStreptomyces phaeoluteichromatogenesStreptomyces polychromogenesStreptomyces racemochromogenesStreptomyces sparsogenesStreptomyces violaceochromogenesStreptomyces viridochromogenesStreptomyces xanthochromogenesStreptosporangium amethystogenesStreptosporangium violaceochromogenesStreptoverticillium parvisporogenes corrig.Suttonella indologenesTepidibacter formicigenesThermoanaerobacterium thermosulfurigenesThermosulfuriphilus ammonigenesToxothrix trichogenesTractidigestivibacter scatoligenesTrichlorobacter thiogenesTrueperella pyogenesVibrio aerogenesVibrio aphrogenesVibrio gazogenesVibrio succinogenesWolinella succinogenes

gignens

"Acetanaerobacter ethanolgignens"Acetivibrio ethanolgignensCaldanaerovirga acetigignensDehalogenimonas alkenigignensThermoanaerobacter sulfurigignens

glaucescens

"Actinomyces glaucescens"Streptomyces glaucescens

glomerans

Coriobacterium glomerans

gregans

Methylobacterium gregans

habitans

Flavimonas oryzihabitansGlycomyces phytohabitansKibdelosporangium phytohabitansLactobacillus suantsaiihabitansLevilactobacillus suantsaiihabitansLevilactobacillus suantsaiihabitansMethylobacterium oryzihabitansNocardia rhizosphaerihabitansPaenibacillus phytohabitansPseudomonas oryzihabitans"Rhizobium oryzihabitans"Sediminibacterium magnilacihabitansStreptomyces phytohabitansStreptomyces rhizosphaerihabitansVibrionimonas magnilacihabitans

hydrogenans

Streptomyces hydrogenans

imitans

Corynebacterium imitansMycoplasma imitans"Mycoplasmoides imitans"

indigens

Azoarcus indigens

inhibens

Carnobacterium inhibensPhaeobacter inhibensRoseovarius nubinhibensSerratia inhibensStreptomyces inhibensVibrio inhibens

interrogans

Leptospira interrogans"Spirochaeta interrogans"

labens

Flavobacterium circumlabens

latens

Burkholderia latens

lineans

"Bacterium albilineans""Phytomonas rubrilineans"Pseudomonas rubrilineansXanthomonas albilineans

linens

"Bacterium linens"Brevibacterium linens

luminescens

Photorhabdus luminescensXenorhabdus luminescens

lutescens

Brevibacterium lutescens"Flavobacterium lutescens"Halomonas lutescensLyngbya lutescensSchizosiphon lutescens"Streptomyces luteolutescens"Streptostemon lutescens

lysans

Gemella haemolysansGemella parahaemolysans"Neisseria haemolysans"

macerans

"Aerobacillus macerans"Bacillus maceransPaenibacillus macerans

mandens

Desulfallas sapomandensDesulfotomaculum sapomandensPetrobacter succinatimandensTepidiphilus succinatimandens

marcescens

Serratia marcescens

miscens

Lactovum miscens

movens

Methanobacterium movens

mutans

Alcanivorax xenomutansAnaerovorax odorimutansAnseongella ginsenosidimutansCaedibacter pseudomutansCaldimicrobium thiodismutansChryseobacterium ginsenosidimutansCompanilactobacillus ginsenosidimutansDesulfolutivibrio sulfodismutansDesulfonatronospira thiodismutansDesulfonatronovibrio thiodismutansDesulfonatronum thiodismutansDesulfovibrio sulfodismutansFlavisolibacter ginsenosidimutansFlavobacterium ginsenosidimutans"Hankyongella ginsenosidimutans""Lactobacillus ginsenosidimutans""Novosphingobium ginsenosidimutans"Olivibacter ginsenosidimutansPedobacter ginsenosidimutansPedococcus ginsenosidimutansPhnomibacter ginsenosidimutansPhycicoccus ginsenosidimutansPseudobacter ginsenosidimutansPseudoflavitalea ginsenosidimutansPseudomonas chloritidismutansRamlibacter ginsenosidimutansRhodanobacter ginsenosidimutansSolirubrobacter ginsenosidimutansSphingobacterium ginsenosidimutansSphingomonas ginsenosidimutansSphingosinicella ginsenosidimutansStreptococcus mutansSyntrophococcus sucromutansTerrabacter ginsenosidimutansThermosulfurimonas dismutans

mysorens

Arthrobacter mysorensGlutamicibacter mysorens