Species names derived from present participles A-M

General remarks

Species epithets whose last component is a present participle (usually ending in -ens or -ans) can be in any gender. They are treated like Group B adjectives and belong to the third declension. The actual gender is determined by the gender of the genus name [Rule 12c (1)]. One part of the list of epithets formed in that manner is provided below. Genus names and subspecies names derived from present participles are not listed here. A special case is genes, which originally is an Ancient Greek suffix meaning "born from" but is nowadays mainly used as a Neolatin suffix meaning "producing". See also the LPSN introductory page on forming names.

New combinations

In the case of a ☞ new combination these epithets must not be adapted to the gender of the genus name. This is a special case of Group B adjectives.

captans

Gluconacetobacter azotocaptans

cohabitans

Rectinema cohabitans

colens

Achromobacter mucicolens — Actinomyces denticolens — Alloscardovia omnicolens — Bifidobacterium denticolens — Corynebacterium accolens — Pandoraea horticolens — Parascardovia denticolens — Pyramidobacter piscolens — Treponema isoptericolens

comitans

Actinobacillus actinomycetemcomitans — Aggregatibacter actinomycetemcomitans — Haemophilus actinomycetemcomitans — Roseivirga echinicomitans — Vibrio comitans

commovens

Pandoraea commovens

contaminans

Alicyclobacillus contaminans — Anoxybacillus contaminans — Burkholderia contaminans — Chryseobacterium contaminans — Herminiimonas contaminans — Lysinibacillus contaminans — Ornithinibacillus contaminans — Paenibacillus contaminans — Paracoccus contaminans — Pseudochelatococcus contaminans — Sphingopyxis contaminans — "Sphingorhabdus contaminans" — Sphingorhabdus contaminans — Sporosarcina contaminans

convertens

Slackia isoflavoniconvertens — Sphingobium aromaticiconvertens

crescens

Aquimarina celericrescens — Arthrobacter celericrescens — Bacillus crescens — Clostridium celerecrescens — Cohaesibacter celericrescens — Kluyvera cryocrescens — Lacrimispora celerecrescens — Liberibacter crescens — Marinobacter segnicrescens — Methylophaga nitratireducenticrescens — Oryzihumus leptocrescens — Spiroplasma velocicrescens — Thermococcus celericrescens — Thiobacillus rapidicrescens

cumulans

Acidiplasma cupricumulans — Acinetobacter cumulans — Catenisphaera adipataccumulans — Ferroplasma cupricumulans — Variovorax boronicumulans

delens

Nocardia cummidelens — Nonlabens xylanidelens — Persicivirga xylanidelens

denitrificans

Bacillus halodenitrificans — Bacillus thermodenitrificans — Geobacillus thermodenitrificans — Halomonas halodenitrificans — "Micrococcus halodenitrificans" — Microvirgula aerodenitrificans — Paracoccus halodenitrificans — Thioalkalivibrio thiocyanodenitrificans corrig. — Virgibacillus halodenitrificans

devorans

"Bacillus devorans" — Flavobacterium devorans — Paenibacillus agaridevorans — Thalassobacillus devorans

dolens

Achromobacter dolens — Pseudomonas mucidolens

durans

Acidithiobacillus ferridurans — Acidocella aluminiidurans — Alkalihalobacillus halodurans — Alkalilimnicola halodurans corrig. — Alsobacter metallidurans — Aridibacter famidurans — Arthrobacter halodurans — Bacillus halodurans — Bacillus psychrodurans — Bacillus salidurans — Bacillus sporothermodurans — Companilactobacillus halodurans — Cupriavidus metallidurans — Deinococcus radiodurans — Desulfosporosinus acididurans — Dyella halodurans — Dyella psychrodurans — Enterococcus durans — Flavobacterium acidurans — Halomonas halodurans — Halomonas zincidurans — Halorubrum halodurans — Lactobacillus halodurans — Lentibacillus halodurans — Maribacter cobaltidurans — Marivibrio halodurans — Mesorhizobium metallidurans — Methanobrevibacter acididurans — Methylobacterium durans — Microbacterium radiodurans — Micromonospora nickelidurans — Nocardioides euryhalodurans — Paenibacillus xerothermodurans — Pediococcus ethanolidurans — Psychrobacillus psychrodurans — Ralstonia metallidurans — Rhizobium metallidurans — Roseivivax halodurans — Roseomonas radiodurans — Salinicoccus halodurans — Streptococcus durans — Thalassococcus halodurans — Wautersia metallidurans

edens

Burkholderia peredens — Caballeronia peredens — Caedibacter varicaedens — "Caedimonas varicaedens" — Defluviitalea raffinosedens — Halorubrum glutamatedens — Williamsia serinedens

egens

Altererythrobacter salegens — Arenibacter catalasegens — Arthrobacter terregens — "Ascidiaceibacter salegens" — Aureobacterium terregens — Beneckea natriegens — Croceibacterium salegens — Flavobacterium salegens — Hallella seregens — Microbacterium terregens — Pseudidiomarina gelatinasegens — "Pseudomonas natriegens" — Pseudomonas protegens — Pseudomonas salegens — Salegentibacter salegens — Sneathia sanguinegens — Vibrio natriegens

exedens

Advenella alkanexedens — "Arenimonas alkanexedens" — Dehalogenimonas formicexedens — Desulfoglaeba alkanexedens — Nannocystis exedens — Paenibacillus agarexedens — Paenibacillus xylanexedens

exigens

Aestuariibacter salexigens — Bacillus salexigens — Bryantella formatexigens — Chromatium salexigens — Chromohalobacter salexigens — Desulfocapsa sulfexigens corrig. — Desulfovibrio salexigens — Desulfuromonas acetexigens — Gramella salexigens — Haliea salexigens — Halochromatium salexigens — Marinobacter salexigens — Marinobacter salinexigens — Marvinbryantia formatexigens — Melitea salexigens — Mycoavidus cysteinexigens — Pararhodospirillum sulfurexigens — "Rheinheimera salexigens" — Rheinheimera salexigens — Rhodopseudomonas pentothenatexigens — Rhodospirillum salexigens — Rhodospirillum sulfurexigens — Rhodothalassium salexigens — Salibacillus salexigens — Salimicrobium salexigens — Thalassobaculum salexigens — Virgibacillus salexigens

faciens

Acetobacter liquefaciens — Acetobacter liquefaciens — Achromobacter aegrifaciens — Acidisphaera rubrifaciens — Adlercreutzia equolifaciens — Agrobacterium tumefaciens — Alteromonas nigrifaciens — Alteromonas putrefaciens — Aquibacter zeaxanthinifaciens — Arthrobacter ureafaciens — Aureobacterium liquefaciens — Bacillus acidinfaciens — Bacillus amyloliquefaciens — Bacillus pulvifaciens — "Bacillus putrefaciens" — Bacteroides acidifaciens — "Bacteroides aerofaciens" — Bacteroides propionicifaciens — Brenneria rubrifaciens — Brevibacterium liquefaciens — Clostridium putrefaciens — Collinsella aerofaciens — Corynebacterium flaccumfaciens — Corynebacterium liquefaciens — Corynebacterium mucifaciens — "Corynebacterium ureafaciens" — Cosenzaea myxofaciens — Curtobacterium flaccumfaciens — Dialister propionicifaciens — Dyella amyloliquefaciens — Ellagibacter isourolithinifaciens — Erwinia rubrifaciens — Eubacterium aerofaciens — Globicatella sulfidifaciens — Gluconacetobacter liquefaciens corrig. — Gordonibacter urolithinfaciens — Kitasatospora aureofaciens — Lactobacillus kefiranofaciens — Lentibacillus amyloliquefaciens — Mesoflavibacter zeaxanthinifaciens — Microbacterium liquefaciens — Micromonospora viridifaciens — Moraxella nonliquefaciens — Mycoplasma lipofaciens — Mycoplasma putrefaciens — Mycoplasmopsis lipofaciens — Natronorubrum sulfidifaciens — Nubsella zeaxanthinifaciens — Paenarthrobacter ureafaciens — Paenibacillus pulvifaciens — Parabacteroides acidifaciens — Paracoccus carotinifaciens — Paracoccus zeaxanthinifaciens — "Pedobacter zeaxanthinifaciens" — Propionibacterium acidifaciens — Proteus myxofaciens — Providencia alcalifaciens — Pseudoalteromonas nigrifaciens — Pseudomonas aureofaciens — Rhizomonas suberifaciens — Rhizorhapis suberifaciens — Robiginitalea myxolifaciens — Rubritalea squalenifaciens — Ruminococcus flavefaciens — Salinicoccus halitifaciens — Serratia liquefaciens — Shewanella putrefaciens — Siansivirga zeaxanthinifaciens — Slackia equolifaciens — Sphingobium suberifaciens — Sphingomicrobium astaxanthinifaciens — Sphingomonas astaxanthinifaciens — Sphingomonas carotinifaciens — Sphingomonas floccifaciens — Sphingomonas spermidinifaciens — Sphingomonas suberifaciens — Streptomyces albofaciens — Streptomyces ambofaciens — Streptomyces aureofaciens — Streptomyces graminofaciens — Streptomyces melanosporofaciens — Streptomyces nigrifaciens — Streptomyces phaeofaciens — Streptomyces roseifaciens — Streptomyces sulfonofaciens — Sulfobacillus benefaciens — Thermus amyloliquefaciens — Treponema succinifaciens

fermentans

"Abyssisolibacter fermentans" — Acetobacter oryzifermentans — Acidaminococcus fermentans — Actinotalea fermentans — Alkalibacter saccharofermentans — Alkaliphilus peptidifermentans corrig. — Anaerotignum lactatifermentans — Anoxybacter fermentans — "Bacillus bifermentans" — Bacillus glycinifermentans — Brachybacterium tyrofermentans — "Brevefilum fermentans" — Brevibacterium fermentans — Caproicibacter fermentans — Cellulomonas fermentans — Clostridium bifermentans — "Clostridium cellulofermentans" — Clostridium lactatifermentans — Clostridium phytofermentans — Corynebacterium afermentans — Cytophaga fermentans — Desulfurococcus fermentans — Dyadobacter fermentans — Gallibacterium trehalosifermentans — Geothrix fermentans — Gluconacetobacter sucrofermentans — Halanaerobium fermentans corrig. — "Komagatabacter sucrofermentans" — Komagataeibacter sucrofermentans — "Lachnoclostridium phytofermentans" — Lactiplantibacillus fabifermentans — Lactobacillus bifermentans — Lactobacillus fabifermentans — Lactobacillus malefermentans — Lactobacillus oligofermentans — Loigolactobacillus bifermentans — Macellibacteroides fermentans — Mycoplasma fermentans — Mycoplasmopsis fermentans — Paludibaculum fermentans — Paraclostridium bifermentans — Paucilactobacillus oligofermentans — Pelosinus fermentans — Proteiniphilum saccharofermentans — "Candidatus Pseudoramibacter fermentans" — "Psychrosinus fermentans" — Rhodoferax fermentans — Rodentibacter trehalosifermentans — Roseiarcus fermentans — Saccharicrinis fermentans — Secundilactobacillus malefermentans — Staphylococcus piscifermentans — Streptococcus oligofermentans — Tepidibacillus fermentans — Vagococcus acidifermentans

figens

Acetobacter nitrogenifigens — Ahniella affigens — Ciceribacter azotifigens — Flavobacterium nitrogenifigens — Ideonella azotifigens — Natronobacillus azotifigens — Novosphingobium nitrogenifigens — Oleiliquidispirillum nitrogeniifigens — Paenibacillus azotifigens — Paraburkholderia azotifigens — Pseudomonas azotifigens — Sphingomonas azotifigens

flagrans

Arvibacter flaviflagrans — Flavobacterium flaviflagrans

fluorescens

Actinomadura luteofluorescens — "Actinomyces citreofluorescens" — Streptomyces citreofluorescens — "Streptomyces luteofluorescens"

formans

Acetomicrobium hydrogeniformans — Acidaminobacter hydrogenoformans — Actinomadura sporangiiformans — Allisonella histaminiformans — Anaerobaculum hydrogeniformans — Bacillus azotoformans — Bacillus ectoiniformans — "Bacillus gonidiaformans" — Bacteroides zoogleoformans — Caecibacterium sporoformans — "Caecibacterium sporoformans" — Capsularis zoogleoformans corrig. — Carboxydothermus hydrogenoformans — Cereibacter azotoformans — Clostridium hydrogeniformans — "Corynebacterium laevaniformans" — Dulcicalothrix necridiiformans — Faecalicoccus acidiformans — Fortiea necridiiformans — Fusobacterium gonidiaformans — "Luteovulum azotoformans" — Meiothermus cateniformans — Microbacterium laevaniformans — Nocardiopsis synnemataformans — Prevotella zoogleoformans — Proteinivorax hydrogeniformans — Pseudomonas azotoformans — Rhizobium rosettiformans — Rhodobacter azotoformans — Ruthenibacterium lactatiformans — Streptomyces sporangiiformans — Streptomyces synnematoformans — Suicoccus acidiformans — Thermoanaerobacterium butyriciformans — Thermosipho globiformans

fugiens

"Candidatus Haliscomenobacter calcifugiens"

genens

Alkalispirochaeta sphaeroplastigenens — Mechercharimyces asporophorigenens — Sphingomonas sanxanigenens — Spirochaeta sphaeroplastigenens — Virgisporangium aliadipatigenens

generans

Dorea formicigenerans — Eubacterium formicigenerans

-genes (genes)

Acetobacteroides hydrogenigenes — Actinobacillus succinogenes — Actinomadura nitritigenes — "Actinomyces diastatochromogenes" — Actinomyces pyogenes — Actinomyces pyogenes — "Actinomyces violaceochromogenes" — "Actinomyces viridochromogenes" — Actinoplanes ianthinogenes — Aeromonas enteropelogenes — Agrobacterium rhizogenes — Alkalibaculum sporogenes — "Anaerophilus nitritogenes" — Aquitalea pelogenes — Arcanobacterium pyogenes — Arthrobacter polychromogenes — Azomonas macrocytogenes — Azomonotrichon macrocytogenes — Azotobacter macrocytogenes — "Bacillus necrogenes" — "Bacillus pyogenes" — "Bacillus scatologenes" — "Bacillus sporogenes" — Bacteroides helcogenes — Bacteroides pyogenes — Bacteroides succinogenes — Beneckea gazogenes — Biomaibacter acetigenes — Brevibacterium ammoniagenes — Caminicella sporogenes — Chryseobacterium indologenes — Clostridium gasigenes — Clostridium nitritogenes — Clostridium scatologenes — Clostridium sporogenes — Clostridium sulfidigenes — Clostridium thermosuccinogenes — Clostridium thermosulfurigenes corrig. — Corynebacterium ammoniagenes — Corynebacterium pyogenes — Curtobacterium ammoniigenes — Desulfocapsa thiozymogenes — Desulfonispora thiosulfatigenes — Enterobacter aerogenes — Ercella succinigenes — Eubacterium coprostanoligenes — Eubacterium nitritogenes — Fibrobacter succinogenes — Flavobacterium indologenes — Fusobacterium necrogenes — Geobacter thiogenes — Halopelagius fulvigenes — Haloplanus aerogenes — Kingella indologenes — Klebsiella aerogenes — Lechevalieria aerocolonigenes — Lentzea aerocolonigenes — "Leptothrix trichogenes" — Listeria monocytogenes — Lysobacter enzymogenes — Mediterraneibacter butyricigenes — Methylobacillus glycogenes — Microbispora amethystogenes — Microbispora chromogenes — Micromonospora purpureochromogenes — Moryella indoligenes — Myceligenerans crystallogenes — Mycobacterium farcinogenes — Mycolicibacterium farcinogenes — "Necromonas achromogenes" — "Nitrosopumilus cobalaminigenes" — "Nocardia aerocolonigenes" — Nocardiopsis chromatogenes — Olsenella scatoligenes — Oscillibacter valericigenes — Oxalobacter formigenes — Paludibacter propionicigenes — Pasteurella aerogenes — Pelobacter seleniigenes — "Phytomonas rhizogenes" — Proteiniborus ethanoligenes — Proteiniphilum acetatigenes — Pseudarthrobacter polychromogenes — "Pseudoclostridium thermosuccinogenes" — Pseudomonas alcaligenes — Pseudomonas pseudoalcaligenes — Psychrobacter fulvigenes — Pyrinomonas methylaliphatogenes — Rhizobium rhizogenes — Rhizobium tumorigenes — Saccharofermentans acetigenes — Saccharothrix aerocolonigenes — Sporanaerobacter acetigenes — Staphylococcus chromogenes — Streptacidiphilus melanogenes — Streptococcus pyogenes — Streptomyces achromogenes — Streptomyces albidochromogenes — Streptomyces cinerochromogenes — Streptomyces cyaneochromogenes — Streptomyces diastatochromogenes — Streptomyces fuscichromogenes — Streptomyces fuscigenes — Streptomyces griseochromogenes — Streptomyces indoligenes — Streptomyces melanogenes — Streptomyces olivochromogenes — Streptomyces parvisporogenes — Streptomyces phaeochromogenes — Streptomyces phaeogriseichromatogenes — Streptomyces phaeoluteichromatogenes — Streptomyces polychromogenes — Streptomyces racemochromogenes — Streptomyces sparsogenes — Streptomyces violaceochromogenes — Streptomyces viridochromogenes — Streptomyces xanthochromogenes — Streptosporangium amethystogenes — Streptosporangium violaceochromogenes — Streptoverticillium parvisporogenes corrig. — Suttonella indologenes — Tepidibacter formicigenes — Thermoanaerobacterium thermosulfurigenes — Thermosulfuriphilus ammonigenes — Toxothrix trichogenes — Trichlorobacter thiogenes — Trueperella pyogenes — Vibrio aerogenes — Vibrio aphrogenes — Vibrio gazogenes — Vibrio succinogenes — Wolinella succinogenes

gignens

Acetivibrio ethanolgignens — Caldanaerovirga acetigignens — Dehalogenimonas alkenigignens — Thermoanaerobacter sulfurigignens

habitans

Flavimonas oryzihabitans — Glycomyces phytohabitans — Kibdelosporangium phytohabitans — Lactobacillus suantsaiihabitans — "Levilactobacillus suantsaiihabitans" — Methylobacterium oryzihabitans — Nocardia rhizosphaerihabitans — Pseudomonas oryzihabitans — "Rhizobium oryzihabitans" — Sediminibacterium magnilacihabitans — Streptomyces phytohabitans — Streptomyces rhizosphaerihabitans — Vibrionimonas magnilacihabitans

labens

Flavobacterium circumlabens

lutescens

Brevibacterium lutescens (sic) — Halomonas lutescens

lysans

Gemella haemolysans — Gemella parahaemolysans — "Neisseria haemolysans"

mutans

Alcanivorax xenomutans — Anaerovorax odorimutans — Anseongella ginsenosidimutans — Caedibacter pseudomutans — Caldimicrobium thiodismutans — Chryseobacterium ginsenosidimutans — Companilactobacillus ginsenosidimutans — "Desulfolutivibrio sulfodismutans" — Desulfonatronospira thiodismutans — Desulfonatronovibrio thiodismutans — Desulfonatronum thiodismutans — Desulfovibrio sulfodismutans — Flavisolibacter ginsenosidimutans — Flavobacterium ginsenosidimutans — "Hankyongella ginsenosidimutans" — "Lactobacillus ginsenosidimutans" — "Novosphingobium ginsenosidimutans" — Olivibacter ginsenosidimutans — Pedobacter ginsenosidimutans — Pedococcus ginsenosidimutans — Phycicoccus ginsenosidimutans — Pseudobacter ginsenosidimutans — Pseudoflavitalea ginsenosidimutans — Pseudomonas chloritidismutans — Ramlibacter ginsenosidimutans — Rhodanobacter ginsenosidimutans — Solirubrobacter ginsenosidimutans — Sphingobacterium ginsenosidimutans — Sphingomonas ginsenosidimutans — Sphingosinicella ginsenosidimutans — Streptococcus mutans — Syntrophococcus sucromutans — Terrabacter ginsenosidimutans — Thermosulfurimonas dismutans