Subspecies and species names using Group B adjectives, part 1

General remarks

Epithets of species or subspecies can be formed from simple or compound Latin or Neolatin adjectives [Rule 12c (1), Rule 13b]. Group B adjectives are declined according to the third declension and either end in -is (masculine), -is (feminine), -e (neuter) or in -er (masculine), -ris (feminine), -re (neuter) or are the same for all genders. One part of the species and subspecies names found in LPSN and derived in that manner are shown here, arranged according to the stem of the adjective. Compound adjectives are grouped according to the last part (the one that is modified according to the gender) as far as possible. See also the LPSN introductory page on forming names.

New combinations

In the case of a new combination these epithets must be adapted to the gender of the genus name: -is (masculine), -is (feminine) or -e (neuter).

affin- (affinis, affinis, affine)

Shewanella affinis

agil- (agilis, agilis, agile)

Achromobacter agilisAnaerofilum agileArthrobacter agilisAzomonas agilis"Azotobacter agile"Lactobacillus agilisLigilactobacillus agilisMassilia agilisMethylobacter agilisMethylomicrobium agileMicrococcus agilisXanthobacter agilis

agrest- (agrestis, agrestis, agreste)

Buttiauxella agrestisThermocrispum agreste

amical- (amicalis, amicalis, amicale)

Gordonia amicalis

aquatil- (aquatilis, aquatilis, aquatile)

Acidibrevibacterium fodinaquatileAlcaligenes aquatilisAlgoriphagus aquatilisAlicyclobacillus fodiniaquatilisAllopseudarcicella aquatilisAmpullimonas aquatilisAquiflexum aquatile"Bacillus aquatilis"Catellibacterium aquatileCellulosimicrobium aquatileChitinilyticum aquatileComamonas aquatilisCytophaga aquatilisDeinococcus aquatilisDuganella aquatilisEmticicia aquatilisFlavobacterium aquatileFluviibacterium aquatileGemmobacter aquatilisHerminiimonas aquatilisHymenobacter aquatilisLacibacterium aquatileLelliottia aquatilisNevskia aquatilis"Paucibacter aquatile"Pedobacter aquatilisPsychromonas aquatilisRahnella aquatilisReyranella aquatilisRoseateles aquatilisSerratia aquatilisSphingomonas aquatilisSynechocystis aquatilisUndibacterium aquatileZavarzinia aquatilis

arval- (arvalis, arvalis, arvale)

Conexibacter arvalis

auricular- (auricularis, auricularis, auriculare)

Staphylococcus auricularis

austral- (australis, australis, australe)

Acidiferrimicrobium australe"Bartonella australis""Burkholderia australis"Glaesserella australisLimnohabitans australis"Magnetofaba australis""Candidatus Neoehrlichia australis"Photorhabdus australisPseudomonas extremaustralisRickettsia australisStreptococcus australisPhotorhabdus asymbiotica subsp. australis

balnear- (balnearis, balnearis, balneare)

Brevundimonas balnearis

boreal- (borealis, borealis, boreale)

Acidicapsa borealisHalomonas borealisPaenibacillus borealisPaludisphaera borealisPedobacter borealisRosenbergiella australiborealis corrig.Spongiibacter borealisZhongshania borealis

brev- (brevis, brevis, breve)

Alkalilacustris brevisBacillus brevisBacillus parabrevisBifidobacterium breveBrevibacillus brevisBrevibacillus parabrevisCryobacterium breveDesertihabitans brevisEmpedobacter brevisFlavobacterium breveHalorientalis brevisJanibacter brevisLactobacillus brevisLactobacillus parabrevisLevilactobacillus brevisLevilactobacillus parabrevisMarinifilum breveMillisia brevisNatribaculum breveOscillatoria brevisOscillatoria subbrevisPontibacter brevisPrevotella brevis"Candidatus Promineofilum breve"Sulfitobacter brevisThermomonas brevisBacteroides ruminicola subsp. brevisPrevotella ruminicola subsp. brevis

caelest- (caelestis, caelestis, caeleste)

Streptomyces caelestis

campestr- (campestris, campestris, campestre)

"Bacillus campestris"Brochothrix campestrisXanthomonas campestris

central- (centralis, centralis, centrale)

Anaplasma centrale

cloacal- (cloacalis, cloacalis, cloacale)

Atopostipes suicloacalis corrig.Metamycoplasma cloacaleMycoplasma cloacale

columnar- (columnaris, columnaris, columnare)

Cytophaga columnarisFlavobacterium columnareFlexibacter columnaris

commun- (communis, communis, commune)

Alistipes communisAlteromonas communisAquabacterium communeAzoarcus communisBifidobacterium communeFlavobacterium communeMarinomonas communisNitrosomonas communisOceanospirillum communePandoraea communis"Candidatus Parabeggiatoa communis"Paracoccus communis"Candidatus Pelagibacter communis" corrig.Thermodesulfobacterium communeVibrio communis

debil- (debilis, debilis, debile)

Caldibacillus debilisGeobacillus debilis

difficil- (difficilis, difficilis, difficile)

Alkanibacter difficilis"Bacillus difficilis"Clostridioides difficileClostridium difficileMethylosoma difficile"Candidatus Planktoluna difficilis"Polynucleobacter difficilisSphingomonas difficilisStreptococcus difficile (sic)Streptococcus difficilis corrig.

exil- (exilis, exilis, exile)

Caldisericum exileWinogradskyella exilis

facil- (facilis, facilis, facile)

Acidiphilium facile corrig.Acidiphilium facilis (sic)Acidocella facilisAcidovorax facilisAlteromonas facilis"Hydrogenomonas facilis"Hyphomicrobium facile corrig.Hyphomicrobium facilis (sic)Pseudomonas facilisVogesella facilisHyphomicrobium facile subsp. facile corrig.Hyphomicrobium facilis subsp. facilis (sic)

fidel- (fidelis, fidelis, fidele)

Shewanella fidelis

flexibil- (flexibilis, flexibilis, flexibile)

Pseudomonas flexibilisSerpens flexibilis

flexil- (flexilis, flexilis, flexile)

Flexibacter flexilisMethanobacterium flexileThiofilum flexileThiothrix flexilis

fluvial- (fluvialis, fluvialis, fluviale)

Aeromonas fluvialisPseudomonas fluvialisRaineyella fluvialisSphingobium fluvialeSpirosoma fluvialeVagococcus fluvialisVibrio fluvialis

fluviatil- (fluviatilis, fluviatilis, fluviatile)

Chromobacterium fluviatileFlavobacterium fluviatileIodobacter fluviatile (sic)Iodobacter fluviatilis corrig.Prosthecobacter fluviatilis

fort- (fortis, fortis, forte)

Alteromonas fortisBacillus fortisBrevibacillus fortisVibrio fortis

fragil- (fragilis, fragilis, fragile)

"Bacillus fragilis"Bacteroides fragilisDesulfofrigus fragileHassallia fragilisLeptolyngbya fragilisMarinifilum fragilePhormidium fragileStreptomyces fragilisStreptosporangium fragileTolypothrix fragilisYoungiibacter fragilis

glacial- (glacialis, glacialis, glaciale)

Arthrobacter glacialisBrumimicrobium glacialeDevosia glacialisHymenobacter glacialisPaenibacillus glacialisPedobacter glacialisPolaromonas glacialisPolymorphobacter glacialisRufibacter glacialisSphingomonas glacialis"Candidatus Vermiplasma glacialis"

gracil- (gracilis, gracilis, gracile)

Alteromonas gracilisAphanizomenon gracileAquaspirillum gracileAquimarina gracilisBacteroides gracilisCampylobacter gracilisChromatium gracileCurvibacter gracilisCyanobium gracileCystobacter gracilisDesulfovibrio gracilisHylemonella gracilisMarichromatium gracileMethylocaldum gracileMucilaginibacter gracilisNitrospina gracilis"Spirillum gracile"Ventosimonas gracilis

grand- (grandis, grandis, grande)

Deinobacter grandisDeinococcus grandisFalcivibrio grandisSaprospira grandis

hortens- (hortensis, hortensis, hortense)

Dechloromonas hortensisOrnithinicoccus hortensis

hospital- (hospitalis, hospitalis, hospitale)

Ignicoccus hospitalis

imperial- (imperialis, imperialis, imperiale)

Brevibacterium imperialeMicrobacterium imperiale

insign- (insignis, insignis, insigne)

Azomonas insignis"Azotobacter insigne"Filomicrobium insignePolychlamydum insigne

irregular- (irregularis, irregularis, irregulare)

Asaccharospora irregularis"Bacillus irregularis"Clostridium irregulare corrig.Clostridium irregularis (sic)"Candidatus Endowatersipora irregularis" corrig.Frankia irregularisHaloglomus irregulare

lev- (levis, levis, leve)

Duganella levisStreptomyces levis

litoral- (litoralis, litoralis, litorale)

Aestuariibacter litoralisAestuariisphingobium litoraleAestuariivirga litoralisAlgoriphagus litoralisAliiglaciecola litoralis"Aquimarina litoralis""Aurantimonas litoralis"Bacillus litoralisBacteriovorax litoralisBauldia litoralis corrig.Bernardetia litoralisClostridium litoraleCobetia litoralisCongregibacter litoralisCysteiniphilum litoraleDesulfovibrio litoralisElstera litoralisErythrobacter litoralisFlexibacter litoralisHalobacillus litoralisHalobacteriovorax litoralisHenriciella litoralisJaaginema litoraleLimnobacter litoralisLujinxingia litoralisLutibacter litoralisLutimaribacter litoralisMaribacter litoralisMarinicella litoralisMarinitoga litoralisMarinobacterium litoraleMarinobacter litoralisMetabacillus litoralisMycobacterium litoraleMycolicibacterium litoraleNocardiopsis litoralisOceanisphaera litoralisOceanispirochaeta litoralisPelagibius litoralisPelagicoccus litoralisPelagicola litoralisPeptoclostridium litoralePigmentiphaga litoralisPontibacillus litoralisPonticoccus litoralisPorticoccus litoralisPseudomonas litoralisRoseobacter litoralisShewanella litoralisSnowella litoralisSpirochaeta litoralisSpiroplasma litoraleStreptomyces litoralisSulfitobacter litoralisThalassobius litoralisThermaerobacter litoralisThermococcus litoralisThiocapsa litoralisUlvibacter litoralisVibrio litoralisVirgibacillus litoralis

mirabil- (mirabilis, mirabilis, mirabile)

Caulobacter mirabilisCyanophanon mirabileKribbella mirabilisLautropia mirabilisProteus mirabilisScytonema mirabileStreptomyces mirabilisSulfurococcus mirabilis

mit- (mitis, mitis, mite)

Streptococcus mitis

mobil- (mobilis, mobilis, mobile)

Acetomicrobium mobile"Achromatium mobile"Alcanivorax mobilisAlkalispirillum mobileAminobacterium mobileAnaerobaculum mobileAnaerosporobacter mobilisAzospirillum largimobile corrig.Bacillus mobilisBeijerinckia mobilisCarnobacterium mobileConglomeromonas largomobilisCroceicoccus mobilisDesulfurococcus mobilisDokdonella immobilisDongia mobilisDyella mobilisEctothiorhodospira mobilisEpibacterium mobileFrigidibacter mobilisGlaciimonas immobilisGulbenkiania mobilisHeliobacillus mobilisKlebsiella mobilisLeptothrix mobilisLysobacter mobilisMacromonas mobilisMarinobacter mobilisMaritalea mobilisMesonia mobilisMethanobacterium mobile corrig.Methanobacterium mobilis (sic)Methanomicrobium mobileMethylococcus mobilisMethylotenera mobilisMycoplasma mobileNitrococcus mobilisNocardioides immobilis"Nordella oligomobilis"Paenibacillus mobilisPelagicoccus mobilisPetrotoga mobilisPhenylobacterium immobilePseudomonas paucimobilisPsychrobacter immobilisPyruvatibacter mobilisRuegeria mobilisRufibacter immobilisSphingomonas parapaucimobilisSphingomonas paucimobilisSuccinispira mobilisSulfuriflexus mobilisThermodesulfobacterium mobileThioflavicoccus mobilisTistrella mobilisTritonibacter mobilisTuwongella immobilisZhangella mobilisZymomonas mobilisConglomeromonas largomobilis subsp. largomobilisTritonibacter mobilis subsp. mobilisZymomonas mobilis subsp. mobilis

moll- (mollis, mollis, molle)

Chryseobacterium molleDeinococcus radiomollisPhormidesmis mollePhormidium molle

municipal- (municipalis, municipalis, municipale)

Thermocrispum municipale

mural- (muralis, muralis, murale)

Bacillus muralisBordetella muralisCitricoccus muralisGeorgenia muralisHalomonas muralisJiangella muralisKrasilnikoviella muralisMethylophaga muralis corrig.Microbacterium muraleModestobacter muralisMycobacterium muraleMycolicibacterium muraleOrnithinimicrobium muralePeribacillus muralisPrauserella muralisWilliamsia muralis

mutabil- (mutabilis, mutabilis, mutabile)

"Actinomyces mutabilis"Nocardiopsis mutabilisSaccharothrix mutabilisStreptomyces mutabilisSaccharothrix mutabilis subsp. mutabilis

nexil- (nexilis, nexilis, nexile)

Clostridium nexile"Tyzzerella nexilis"

nival- (nivalis, nivalis, nivale)

Conyzicola nivalisSulfuriferula nivalis

oral- (oralis, oralis, orale)

"Actinomyces oralis"Bacteroides oralisBacteroides veroralisDesulfomicrobium oraleKingella orale (sic)Kingella oralis corrig.Lachnoanaerobaculum oraleMetamycoplasma oraleMethanobrevibacter oralisMycoplasma columboraleMycoplasma oraleMycoplasmopsis columboralisNeisseria oralis"Paenibacillus oralis"Pasteurella oralisPrevotella oralisPrevotella veroralisStreptococcus oralisUreaplasma galloraleStreptococcus oralis subsp. oralis

oriental- (orientalis, orientalis, orientale)

Acetobacter orientalisAmorphus orientalisAmycolatopsis orientalisDraconibacterium orientaleFlavobacterium orientaleFrancisella orientalisGracilibacillus orientalisHalorubrum orientaleMaribacter orientalisNocardia orientalisPseudomonas extremorientalisPseudomonas orientalisRippkaea orientalisSorangium orientale"Streptomyces orientalis"Vibrio orientalisAmycolatopsis orientalis subsp. orientalisFrancisella noatunensis subsp. orientalis

palustr- (palustris, palustris, palustre)

Azospirillum palustreChryseobacterium palustreDesulfocucumis palustrisFlavobacterium palustreKocuria palustrisMethanobacterium palustreMethanosphaerula palustrisMethylocella palustrisMycobacterium palustreRhizomicrobium palustreRhodopseudomonas palustrisRhodopseudomonas parapalustrisRhodopseudomonas pseudopalustrisRuminococcus palustrisRunella palustrisSphingomonas palustrisTrichococcus palustrisVibrio palustris

perenn- (perennis, perennis, perenne)

Clostridium perenne

pratens- (pratensis, pratensis, pratense)

Agreia pratensisCollimonas pratensisLuteitalea pratensisMethylobacillus pratensisSkermanella pratensisStreptomyces pratensisSubtercola pratensis

puteal- (putealis, putealis, puteale)

Desulfovibrio putealisMicrocella putealisSphingobacterium puteale

regular- (regularis, regularis, regulare)

Actinoplanes regularis"Ampullaria regularis"Ampullariella regularisChamaesiphonopsis regularisCoelomoron regularisHalorientalis regularis

rud- (rudis, rudis, rude)

Acinetobacter rudis

saecular- (saecularis, saecularis, saeculare)

Bifidobacterium saeculareBifidobacterium pullorum subsp. saeculare

segn- (segnis, segnis, segne)

Aggregatibacter segnisAquamicrobium segneCaulobacter segnisHaemophilus segnisMycoplana segnis

septentrional- (septentrionalis, septentrionalis, septentrionale)

"Candidatus Cryosericum septentrionale"Mesorhizobium septentrionalePaenibacillus septentrionalis"Streptomyces septentrionalis"

silvestr- (silvestris, silvestris, silvestre)

Bacillus silvestrisLysobacter silvestrisMarmoricola silvestrisMethylocella silvestrisNakamurella silvestrisPontibacter silvestrisPsychromicrobium silvestreSolibacillus silvestrisSolimicrobium silvestreVicinamibacter silvestris

simil- (similis, similis, simile)

Arcanobacterium phocisimileKerstersia similisLactobacillus similisMarinobacter similisSecundilactobacillus similisYersinia similisStreptococcus dysgalactiae subsp. equisimilis

singular- (singularis, singularis, singulare)

Burkholderia singularisCalenema singularisCorynebacterium singulareGlaciimonas singularis"Candidatus Thioglobus singularis"

special- (specialis, specialis, speciale)

Streptomyces specialis

spectabil- (spectabilis, spectabilis, spectabile)

Streptomyces spectabilis

spiral- (spiralis, spiralis, spirale)

Actinomadura spiralisCaedibacter taeniospiralisElytrosporangium spiraleHydrogenophaga taeniospiralisKatagnymene spiralisMicrotetraspora spiralisNonomuraea spiralis corrig."Pseudoalteromonas spiralis"Pseudomonas taeniospiralisStreptomyces capillispiralis"Streptomyces gilvospiralis"Streptomyces spiralis

stabil- (stabilis, stabilis, stabile)

Burkholderia stabilis

subtil- (subtilis, subtilis, subtile)

Ancylomarina subtilisBacillus subtilisBifidobacterium subtilePlanktolyngbya subtilis"Vibrio subtilis"Bacillus subtilis subsp. subtilis

tenu- (tenuis, tenuis, tenue)

"Bacteroides tenuis"Caryophanon tenueEubacterium tenueGlycomyces tenuisHymenobacter tenuisLeptolyngbya tenuisNitrosospira tenuisOscillatoria tenuisPhormidium tenueRhodocyclus tenuisRhodospirillum tenue"Spirochaeta pertenuis"Tolypothrix tenuisTreponema pertenueTrichodesmium tenue"Treponema pallidum subsp. pertenue"

terrestr- (terrestris, terrestris, terrestre)

Alicyclobacillus acidoterrestrisAncylothrix terrestrisBacillus acidoterrestrisBurkholderia terrestrisCaballeronia terrestrisHalobiforma haloterrestrisHalonotius terrestrisHalorubrum terrestreKineococcus terrestris corrig.Kitasatospora terrestrisMongoliimonas terrestris

tumoral- (tumoralis, tumoralis, tumorale)

Pseudonocardia antitumoralis corrig.

universal- (universalis, universalis, universale)

Cronobacter universalisMethyloversatilis universalis

util- (utilis, utilis, utile)

Staphylococcus carnosus subsp. utilis

variabil- (variabilis, variabilis, variabile)

Acinetobacter variabilis"Actinomyces variabilis"Anabaena variabilisArthrobacter variabilisBrevundimonas variabilisCaulobacter variabilisCellulosimicrobium variabileCorynebacterium variabile corrig.Corynebacterium variabilis (sic)Cyanogastrum variabileDesulfosarcina variabilisEctothiorhodospira variabilisHalomonas variabilisHalovibrio variabilisIsoptericola variabilisLimimaricola variabilisLoktanella variabilisMethylobacterium variabileMicrobulbifer variabilisStreptomyces flavovariabilis"Streptomyces olivovariabilis"Streptomyces variabilisSubdoligranulum variabileSulfurifustis variabilisTrichormus variabilisVibrio variabilis

versatil- (versatilis, versatilis, versatile)

Aquimarina versatilis"Candidatus Korobacter versatilis" corrig.Methylotenera versatilisPectobacterium versatileSaccharedens versatilis

virid- (viridis, viridis, viride)

Actinomadura viridis"Actinomyces alboviridis""Actinomyces olivaceoviridis""Actinomyces olivoviridis""Actinomyces roseoviridis"Amycolatopsis oliviviridisAmycolatopsis viridisBlastochloris sulfoviridisBlastochloris viridisChitinimonas viridisClostridium virideKitasatospora viridisMicrobispora viridisMicrocystis viridisMicromonospora fulviviridisMicromonospora luteiviridisMicrotetraspora viridisOscillatoria nigroviridis"Promicromonospora viridis"Rhodobacter viridisRhodopseudomonas sulfoviridisRhodopseudomonas viridisRhodovulum virideSaccharomonospora viridisStauromatonema virideStreptomyces alboviridisStreptomyces flavoviridisStreptomyces griseoviridisStreptomyces olivaceoviridisStreptomyces olivoviridisStreptomyces phaeoviridisStreptomyces roseoviridisStreptomyces viridis"Thermoactinomyces viridis"

vital- (vitalis, vitalis, vitale)

Leptolinea tardivitalis

vulgar- (vulgaris, vulgaris, vulgare)

Balneola vulgarisCellvibrio vulgarisDesulfovibrio vulgarisErythrobacter vulgarisExtensimonas vulgarisHyphobacterium vulgareHyphomicrobium vulgareKetogulonicigenium vulgare corrig."Candidatus Maribeggiatoa vulgaris"Marinobacter vulgarisNitrobacter vulgarisProteus vulgarisSalimesophilobacter vulgarisSphingobium vulgareStreptomyces thermovulgarisStreptosporangium pseudovulgareStreptosporangium vulgareTectibacter vulgarisThermoactinomyces vulgaris"Thermomonospora vulgaris"Alistipes onderdonkii subsp. vulgarisDesulfovibrio vulgaris subsp. vulgaris