Subspecies names derived from present participles

General remarks

Subspecies epithets whose last component is a present participle (usually ending in -ens or -ans) can be in any gender. They are treated like Group B adjectives and belong to the third declension. The actual gender is determined by the gender of the genus name [Rule 12c (1), Rule 13b]. A list of epithets formed in that manner is provided below. Genus names and species names derived from present participles are not listed here. A special case is genes, which originally is an Ancient Greek suffix meaning "born from" but is nowadays mainly used as a Neolatin suffix meaning "producing". See also the LPSN introductory page on forming names.

New combinations

In the case of a new combination these epithets must not be adapted to the gender of the genus name. This is a special case of Group B adjectives.


Nocardia salmonicida subsp. cummidelens


Geobacillus thermodenitrificans subsp. thermodenitrificans


"Nannocystis exedens subsp. exedens"


Acetobacter aceti subsp. liquefaciensAdlercreutzia equolifaciens subsp. equolifaciensBacillus amyloliquefaciens subsp. amyloliquefaciensCorynebacterium flaccumfaciens subsp. flaccumfaciensLactobacillus kefiranofaciens subsp. kefiranofaciensLeucobacter celer subsp. astrifaciensMesoflavibacter zeaxanthinifaciens subsp. zeaxanthinifaciensPaenibacillus larvae subsp. pulvifaciensPseudomonas chlororaphis subsp. aureofaciensStreptomyces griseus subsp. solvifaciens


Acetobacter xylinum subsp. sucrofermentans (sic)Acetobacter xylinus subsp. sucrofermentans corrig.Corynebacterium afermentans subsp. afermentansGluconacetobacter xylinus subsp. sucrofermentansParaclostridium bifermentans subsp. bifermentansStreptomyces cinereoruber subsp. fructofermentans

-genes (genes)

Aeromonas hydrophila subsp. anaerogenesAeromonas salmonicida subsp. achromogenesAzotobacter nigricans subsp. achromogenesBeijerinckia indica subsp. lacticogenesFibrobacter succinogenes subsp. succinogenesGluconobacter oxydans subsp. melanogenesPantoea stewartii subsp. indologenesPseudomonas pseudoalcaligenes subsp. pseudoalcaligenesSaccharothrix aerocolonigenes subsp. aerocolonigenesStaphylococcus hyicus subsp. chromogenesStreptomyces achromogenes subsp. achromogenesStreptosporangium amethystogenes subsp. amethystogenes


Sphaerotilus natans subsp. natans


Achromobacter xylosoxidans subsp. xylosoxidansAlcaligenes xylosoxidans subsp. xylosoxidans


Alcaligenes denitrificans subsp. xylosoxydans"Burkholderia kururiensis subsp. thiooxydans"Gluconobacter oxydans subsp. suboxydans


Geobacter sulfurreducens subsp. sulfurreducensLeucobacter chromiireducens subsp. chromiireducensNeisseria elongata subsp. nitroreducens


Enterobacter cloacae subsp. dissolvens


Hyphomicrobium facile subsp. tolerans corrig.Hyphomicrobium facilis subsp. tolerans (sic)Lacticaseibacillus paracasei subsp. toleransLactobacillus casei subsp. toleransLactobacillus paracasei subsp. tolerans


Geobacillus subterraneus subsp. aromaticivoransPseudomonas oleovorans subsp. oleovoransSphaerotilus natans subsp. sulfidivorans