Name: Shouchella rhizosphaerae (Madhaiyan et al. 2013) Joshi et al. 2022

Category: Species

Proposed as: comb. nov.

Basonym: Bacillus rhizosphaerae Madhaiyan et al. 2013

Etymology: rhi.zo.sphae’rae. N.L. gen. fem. n. rhizosphaerae, of the rhizosphere

Pronunciation (Latin), gender: s͡hou-K^HEL-la r^hi-zo-SP^HA͡E-ra͡e, feminine

Type strain: DSM 21911; NCCB 100267; SC-N012

See detailed strain information at

Conduct genome-based taxonomy at

16S rRNA gene: FJ233848 Analyse FASTA

Effective publication: Joshi A, Thite S, Karodi P, Joseph N, Lodha T. Alkalihalobacterium elongatum gen. nov. sp. nov.: An Antibiotic-Producing Bacterium Isolated From Lonar Lake and Reclassification of the Genus Alkalihalobacillus Into Seven Novel Genera. Front Microbiol 2021; 12:722369. 🇮🇳

IJSEM list: Oren A, Garrity GM. Validation list no. 206. List of new names and new combinations previously effectively, but not validly, published. Int J Syst Evol Microbiol 2022; 72:5422.

Nomenclatural status: validly published under the ICNP

Taxonomic status: synonym (and not recommended for medical use)

Correct name: Shouchella clausii (Nielsen et al. 1995) Joshi et al. 2022

Risk group: 1

• Synonyms:

  • Name	Kind
    Shouchella clausii (Nielsen et al. 1995) Joshi et al. 2022	heterotypic synonym, validly published under the ICNP
    Bacillus clausii Nielsen et al. 1995	heterotypic synonym, validly published under the ICNP
    Bacillus rhizosphaerae Madhaiyan et al. 2013	homotypic synonym, validly published under the ICNP
    Alkalihalobacillus clausii (Nielsen et al. 1995) Patel and Gupta 2020	heterotypic synonym, validly published under the ICNP
    Alkalihalobacillus rhizosphaerae (Madhaiyan et al. 2013) Patel and Gupta 2020	homotypic synonym, validly published under the ICNP

• Notes:
  • ⏲ According to Liu et al. (2019), this species is a later heterotypic synonym of Alkalihalobacillus clausii (Nielsen et al. 1995) Patel and Gupta 2020. Publication: Liu GH, Narsing Rao MP, Dong ZY, Wang JP, Che JM, Chen QQ, Sengonca C, Liu B, Li WJ. Genome-based reclassification of Bacillus plakortidis Borchert et al. 2007 and Bacillus lehensis Ghosh et al. 2007 as a later heterotypic synonym of Bacillus oshimensis Yumoto et al. 2005; Bacillus rhizosphaerae Madhaiyan et al. 2011 as a later heterotypic synonym of Bacillus clausii Nielsen et al. 1995. Antonie Van Leeuwenhoek 2019; 112:1725-1730. 🇨🇳
  • ⏲ According to Liu et al. (2019), this species is a later heterotypic synonym of Bacillus clausii Nielsen et al. 1995. Publication: Liu GH, Narsing Rao MP, Dong ZY, Wang JP, Che JM, Chen QQ, Sengonca C, Liu B, Li WJ. Genome-based reclassification of Bacillus plakortidis Borchert et al. 2007 and Bacillus lehensis Ghosh et al. 2007 as a later heterotypic synonym of Bacillus oshimensis Yumoto et al. 2005; Bacillus rhizosphaerae Madhaiyan et al. 2011 as a later heterotypic synonym of Bacillus clausii Nielsen et al. 1995. Antonie Van Leeuwenhoek 2019; 112:1725-1730. 🇨🇳
  • ⏲ According to Liu et al. (2019), this species is a later heterotypic synonym of Shouchella clausii (Nielsen et al. 1995) Joshi et al. 2022. Publication: Liu GH, Narsing Rao MP, Dong ZY, Wang JP, Che JM, Chen QQ, Sengonca C, Liu B, Li WJ. Genome-based reclassification of Bacillus plakortidis Borchert et al. 2007 and Bacillus lehensis Ghosh et al. 2007 as a later heterotypic synonym of Bacillus oshimensis Yumoto et al. 2005; Bacillus rhizosphaerae Madhaiyan et al. 2011 as a later heterotypic synonym of Bacillus clausii Nielsen et al. 1995. Antonie Van Leeuwenhoek 2019; 112:1725-1730. 🇨🇳
  • 🏢 Synonymy of this taxon was mentioned in an IJSEM list. Publication: Oren A, Garrity GM. List of changes in taxonomic opinion no. 32. Notification of changes in taxonomic opinion previously published outside the IJSEM. Int J Syst Evol Microbiol 2020; 70:4061-4090.
  • ❗ Shouchella rhizosphaerae is the correct name instead if this species is regarded as a separate species (i.e., if its nomenclatural type is not assigned to another species whose name is validly published, legitimate and not rejected and has priority) within a separate genus Shouchella.
  • 🧬 The phylogenomic assignment score of this taxon is 0.28955 (N = 55).
  • 😷 The risk group for Germany has been imported on 2026-08-13. The full classification is: risk group = 1. — If in doubt, use the risk group given in the regulations for your country and, if these are not available, use the risk group given in the catalogue of the culture collection from which you have obtained or intend to obtain the strain.
  • 🎓 Name mentioned 1 time in PubMed until 2026-06-11.
  • 🙅 This name is on the List of Recommended Names for bacteria of medical importance (LoRN) as a synonym of the name to be applied to this species. The name is on LoRN because of the recorded risk group of other species in the genus.

Parent taxon: Shouchella Joshi et al. 2022

Assigned by: Joshi A, Thite S, Karodi P, Joseph N, Lodha T. Alkalihalobacterium elongatum gen. nov. sp. nov.: An Antibiotic-Producing Bacterium Isolated From Lonar Lake and Reclassification of the Genus Alkalihalobacillus Into Seven Novel Genera. Front Microbiol 2021; 12:722369. 🇮🇳

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Record number: 29565